Muzzle Bite
©Monty Sloan
According to the Wolf Ethogram (Wolf Park, Indiana), the Hard Muzzle Bite means “grabbing the muzzle and applying enough force to make the grabbed wolf whimper. Muzzle biting is often accompanied by other threat behaviors which may also elicit whimpering.” Notice the threatening agonistic pucker and lumpy whisker bed of the biting wolf.